Under Construction


Ralph W. Holzenthal, Roger J. Blahnik, Aysha Prather, and Karl Kjer
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
taxon links [up-->]Larcasinae [up-->]Goerinae [up-->]Lepaniinae [down<--]Plenitentoria Interpreting the tree
close box

This tree diagram shows the relationships between several groups of organisms.

The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

example of a tree diagram

You can click on the root to travel down the Tree of Life all the way to the root of all Life, and you can click on the names of descendent subgroups to travel up the Tree of Life all the way to individual species.

For more information on ToL tree formatting, please see Interpreting the Tree or Classification. To learn more about phylogenetic trees, please visit our Phylogenetic Biology pages.

close box
Containing group: Plenitentoria


This is a widely distributed family, found on all continents except South America and Australia. Ulmer (1903) originally described this group as a subfamily of Sericostomatidae. Flint (1960) and other North American workers considered it a subfamily of Limnephilidae, but other authors either always considered it a separate family (Schmid 1980) or elevated the group to its place as a separate family (Wiggins 1996).

Larvae of Goeridae construct cases entirely of rock fragments; some genera incorporate larger rock fragments laterally. Most larvae live in cool running waters and are grazers on periphyton. Lepania larvae are detritivores in spring seepages (Wiggins 1973b). Archithremma ulachensis Martynov is unusual in having a terrestrial pupa (Levanidova & Vshivkova 1984).  (From Holzenthal et al, 2007)


Flint, O.S., Jr. (1960) Taxonomy and biology of Nearctic limnephilid larvae (Trichoptera), with special reference to species in eastern United States. Entomologica Americana, 40, 1–120.

Holzenthal R.W., Blahnik, R.J., Prather, A.L., and Kjer K.M. 2007. Order Trichoptera Kirby 1813 (Insecta), Caddisflies. In: Zhang, Z.-Q., and Shear, W.A. (Eds). 2007 Linneaus Tercentenary: Progress in Invertebrate Taxonomy. Zootaxa. Zootaxa 1668:639–698.

Levanidova, I.M. & Vshivkova, T.S. (1984) The terrestrial pupa of Architremma [Archithremma] ulachensis Mart. (Trichoptera:Limnephilidae); description and considerations. Aquatic Insects, 6, 65–69.

Schmid, F. (1980) Esquisse pour une classification et une phylogénie des Goérides (Trichoptera). Naturaliste Canadien, 107, 185–194.

Ulmer, G. (1903) Ueber die Metamorphose der Trichopteren. Abhandlungen des Naturwissenschaftlichen vereins in Hamburg, 18, 1–154.

Wiggins, G.B. (1973) New systematic data for the North American caddisfly genera Lepania, Goeracea and Goerita (Trichoptera: Limnephilidae). Life Science Contribution, Royal Ontario Museum, 91, 1–33.

Wiggins, G.B. (1996) Larvae of the North American Caddisfly Genera (Trichoptera). University of Toronto Press, Toronto, 457 pp.

Title Illustrations
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Scientific Name Goera pilosa
Specimen Condition Dead Specimen
Identified By Hans Malicky
Life Cycle Stage Adult
View Lateral
Source Collection Barcode of Life Database (BOLD)
About This Page

Ralph W. Holzenthal
University of Minnesota, St. Paul, Minnesota, USA

Roger J. Blahnik
University of Minnesota, St. Paul, Minnesota, USA

Aysha Prather
Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, Toronto, Ontario, Canada

Karl Kjer
Rutgers University, New Brunswick, New Jersey, USA

Correspondence regarding this page should be directed to Ralph W. Holzenthal at , Roger J. Blahnik at , Aysha Prather at , and Karl Kjer at

Page: Tree of Life Goeridae. Authored by Ralph W. Holzenthal, Roger J. Blahnik, Aysha Prather, and Karl Kjer. The TEXT of this page is licensed under the Creative Commons Attribution-NonCommercial-ShareAlike License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Holzenthal, Ralph W., Roger J. Blahnik, Aysha Prather, and Karl Kjer. 2010. Goeridae. Version 20 July 2010 (under construction). http://tolweb.org/Goeridae/14619/2010.07.20 in The Tree of Life Web Project, http://tolweb.org/

edit this page
close box

This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

close box


Page Content

articles & notes



Explore Other Groups

random page

  go to the Tree of Life home page