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Jeff Skevington
Containing group: Pipunculinae


This genus of 274 valid species has a world-wide distribution. Most Tomosvaryella are parasitoids of Cicadellidae.

When looking for information on Palaearctic Tomosvaryella, you may encounter a reference to a review paper by Kuznetzov. Note that the following journal article was never published although it was cited by Kuznetzov in other papers:

Kuznetzov, S. Y. 1995. A review of the Palaearctic genera Tomosvaryella Aczél and Adella gen. n. (Diptera, Pipunculidae). International Journal of Zoological Research 6:177-316.



These stocky little flies are easy to recognize and particularly common in arid or grassland settings. They lack a pterostigma on the wing (within the Pipunculinae only Dorylomorpha and a few species of Eudorylas share this character). They can be differentiated from Dorylomorpha by the shape of the abdomen (widest in the middle). In the few Eudorylini that lack a pterostigma, r-m is in the basal third of the wing and the abdomen is more densely pollinose.

Useful regional revisions include: Afrotropical – De Meyer (1993), Central America, Mexico and the West Indes – Ale-Rocha and Rafael (1995), Central and west European – Földvári and De Meyer (2000), Indian – Kapoor et al. (1987), Japan – Morakote et al. (1990), South America – Ale-Rocha (1996). No recent revisions are available in the Nearctic and Australasian regions and species in these areas are virtually unidentifiable using available keys.



Aczél, M. L. 1939. Die untergattung Dorylomorpha m. von Tomosvaryella m. Dorylaiden-Studien II. Zoologischer Anzeiger 125:49-69.

Aczél, M. L. 1944. Die gattung Tomosvaryella Acz. (Dipt.). (Dorylaiden-Studien VIII). Annales Historico-Naturales Musei Nationalis Hungarici 37:75-130.

Ale-Rocha, R. 1992a. Descriçăo de Tomosvaryella pulchra sp.n. (Diptera: Pipunculidae). Boletim do Museu Paraense Emilio Goeldi, Séries Zoologia 8:315-318.

Ale-Rocha, R. 1992b. Redescriçăo de Tomosvaryella galapagensis (Curran 1934) (Diptera: Pipunculidae). Boletim do Museu Paraense Emilio Goeldi, Séries Zoologia 8:319-321.

Ale-Rocha, R. 1993. Redescriçăo e consideraçőes sobre a distribuiçăo geográfica de Tomosvaryella lepidipes Hardy, 1943 e descriçăo de Tomosvaryella venezuelana, sp. n. (Diptera, Pipunculidae). Revista Brasileira de Entomologia 37:263-266.

Ale-Rocha, R. 1996. Revisăo das espécies de Tomosvaryella Aczél da América do Sul (Diptera, Pipunculidae). [Revision of South American species of Tomosvaryella Aczél (Diptera, Pipunculidae)]. Revista Brasileira de Entomologia 40:165-187.

Ale-Rocha, R., and J. A. Rafael. 1995. Tomosvaryella Aczél do México, América Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia 12:407-427.

Belcari, A. 1994. Note bio-etologiche su Tomosvaryella frontata (Becker) (Diptera: Pipunculidae), parassitoide di Opsius stactogalus Fieber (Homoptera: Cicadellidae). [Bio-ethological notes on Tomosvaryella frontata (Becker) (Diptera: Pipunculidae), a parasitoid of Opsius stactogalus Fieber (Homoptera: Cicadellidae)]. Atti del Congresso Nazionale Italiano di Entomologia 17:573-576.

Belcari, A. 1999. Ethological and morphological notes on Tomosvaryella frontata (Diptera: Pipunculidae), parasitoid of the tamarix leafhopper Opsius stactogalus. Entomological Problems 30:35-46.

De Meyer, M. 1990. A new Tomosvaryella Aczél, 1939 species from Ethiopia (Diptera, Pipunculidae). Bulletin et Annales de la Société Royale belge d'Entomologie 126:87-90.

De Meyer, M. 1993. A revision of the Afrotropical species of Tomosvaryella Aczél, 1939(Diptera: Pipunculidae). Annals of the Natal Museum 34:43-101.

De Meyer, M. 1995. The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana 18:283-319.

De Meyer, M. 1997. Contribution to the Pipunculidae fauna of Spain: (Diptera). Beiträge zur Entomologie 47:421-450.

Földvári, M., and M. De Meyer. 2000. Revision of central and west European Tomosvaryella Aczél species (Diptera, Pipunculidae). Acta Zoologica Academiae Scientiarum Hungaricae 45:299-334.

Hardy, D. E. 1943. A revision of Nearctic Dorilaidae (Pipunculidae). University of Kansas Science Bulletin 29:1-231.

Hardy, D. E. 1958. Proposed use of the plenary powers to suppress the generic names Prothechus Rondani, 1856, and Alloneura Rondani, 1856, for the purpose of validating the generic names Verrallia Mik, 1899, and Tomosvaryella Aczél, 1939 (Class Insecta, Order Diptera). Bulletin of Zoological Nomenclature 17:27-29.

Hardy, D. E. 1967. Redescription of Tomosvaryella frontata (Diptera: Pipunculidae). Annals of the Entomological Society of America 60:116-118.

International Commision on Zoological Nomenclature. 1961. Opinion 597. Prothechus Rondani, 1856, and Alloneura Rondani, 1856, (Insecta, Diptera); suppressed under the Plenary Powers. Bulletin of Zoological Nomenclature 18:230-235.

Kapoor, V. C., J. S. Grewal, and S. K. Sharma. 1987. Indian Pipunculids (Diptera: Pipunculidae). A Comprehensive Monograph. Atlantic Publishers & Distributors, New Delhi, India.

Kehlmaier, C. 2008. Finnish Pipunculidae (Diptera) studies Part I: Taxonomic notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939. Zootaxa 1672: 1-42.

Kirk-Spriggs, A. H. 1988. The female and puparium of Tomosvaryella singula Hardy (Dipt., Pipunculidae). The Entomologist's monthly Magazine 124:181-185.

Kuznetzov, S. Y. 1993a. A new European species of Tomosvaryella (Diptera: Pipunculidae). Zoosystematica Rossica 2:177-178.

Kuznetzov, S. Y. 1993b. New species of Tomosvaryella Aczél (Diptera, Pipunculidae) from Central Asia. Dipterological Research 4:209-224.

Kuznetzov, S. Y. 1993c. New species of Tomosvaryella Aczél (Diptera, Pipunculidae) from Kazakhstan. Dipterological Research 4:163-169.

Kuznetzov, S. Y. 1993d. A new Tomosvaryella Aczél from Egypt, with redescription of T. disjuncta (Becker) (Diptera, Pipunculidae). Dipterological Research 4:151-162.

Kuznetzov, S. Y. 1994a. New Palaearctic species of Tomosvaryella Aczél (Diptera, Pipunculidae) with a redescription of T. rondanii (Collin). Dipterological Research 5:107-207.

Kuznetzov, S. Y. 1994b. A new species of Tomosvaryella Aczél (Diptera, Pipunculidae) from Turkmenistan. Dipterological Research 5:27-31.

Morakote, R., Y. Hirashima, and K. Yano. 1990. A systematic study of the Japanese Pipunculidae (Diptera) Part VI. The genera Dorylomorpha Aczél and Tomosvaryella Aczél. Journal of the Faculty of Agriculture, Kyushu University 34:357-395.

Parker, H. L. 1967. Notes on the biology of Tomosvaryella frontata (Diptera: Pipunculidae), a parasite of the leafhopper Opsius stactogalus on Tamarix. Annals of the Entomological Society of America 60:292-295.

Pierce, D. H. 1972. Scaphytopius nitridus: Parasitization by Tomosvaryella appendipes. Environmental Entomology 1:796-797.

Rafael, J. A. 1996. Pipunculidae (Insecta, Diptera) of the Dominican Republic: new records and description of new species. Annals of the Carnegie Museum 65:363-381.

Rafael, J. A., and M. S. S. Rosa. 1991. Pipunculidae (Diptera) da Estaçăo Ecológica de Maracá e da localidade de Pacaraima, Roraima, Brasil. Acta Amazonica 21:337-350.

Scarbrough, A. G., and L. V. Knutson. 1989. Asilidae, Bombyliidae, Conopidae, and Pipunculidae (Diptera) of Dominica, West Indies. Florida Entomologist 72:519-537.

Skevington, J.H. and Földvári, M. 2007. Revision of Fijian Tomosvaryella Aczél (Diptera: Pipunculidae). In: Evenhuis, N.L. & Bickel, D.J. (eds.), Fiji Arthropods VIII. Bishop Museum Occasional Papers 93: 27-40.

Woodley, N. E., and D. J. Hilburn. 1994. The Diptera of Bermuda. Contributions of the American Entomological Institute 28:1-64.

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Agriculture and Agri-Food Canada, Ottawa, Ontario, Canada

Correspondence regarding this page should be directed to Jeff Skevington at

Page: Tree of Life Tomosvaryella. Authored by Jeff Skevington. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Skevington, Jeff. 2008. Tomosvaryella. Version 07 August 2008 (under construction). http://tolweb.org/Tomosvaryella/54668/2008.08.07 in The Tree of Life Web Project, http://tolweb.org/

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