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Jeff Skevington
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Containing group: Pipunculinae


This is the most speciose genus of pipunculids and contains 415 valid species. The genus is cosmopolitan in distribution and considerable work remains to be done at the species level in many parts of the world. Most Eudorylas are parasitoids of Cicadellidae.


This group of flies can be difficult to recognize at the generic level. Particularly females of Clistoabdominalis and Dasydorylas are difficult to differentiate from some female Eudorylas. Use the key in Skevington and Yeates (2001) to identify Eudorylas.

Useful regional revisions include: Australia – Skevington (2003), Costa Rica – Rafael and Menezes (1999), European – Kehlmaier (2005), Indian – Kapoor et al. (1987), Japan – Morakote et al. (1990), Mexico and Central America – Rafael (1993), and South America – Rafael (1995). No recent revisions are available in the Nearctic, east Palaearctic and Oriental regions and species in these areas are virtually unidentifiable using available resources. A revision of the African species of Eudorylini is expected soon (Földvári, submitted).

Discussion of Phylogenetic Relationships

A tribal phylogeny of the Eudorylini (Skevington and Yeates, 2000) includes several Eudorylas exemplars. A phylogeny of the Australian species is presented in Skevington (2003).


Ackland, D. M. 1999. A new species of Eudorylas from Scotland (Diptera, Pipunculidae). Dipterists Digest 6:35-39.

De Meyer, M. 1995. The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana 18:283-319.

De Meyer, M. 1997. Contribution to the Pipunculidae fauna of Spain: (Diptera). Beiträge zur Entomologie 47:421-450.

De Meyer, M., and J. Skevington. 2001. Eudorylas Aczél, 1940 (Insecta, Diptera): proposed conservation of useage by the designation of Pipunculus fuscipes Zetterstedt, 1844 as the type species. Bulletin of Zoological Nomenclature 58:19-23.

Dempewolf, M. 1996. Taxonomical notes on west palaearctic Pipunculide including the description of two new Eudorylas species (Diptera). Studia dipterologica 3:87-92.

Dempewolf, M. 1998. On the taxonomy of two German pipunculid flies with remarks on the terminology of the male genitalia (Diptera). Beiträge zur Entomologie 48:531-537.

Dempewolf, M., and K. von der Dunk. 1996. A new Eudorylas species from Southern and Western Germany (Diptera, Pipunculidae). Studia dipterologica 3:93-96.

Dunk, K. v. d. 1995. Descriptions of two new Eudorylas species and of the so far unknown female of E. restrictus Coe, 1966 (Dipt.: Pipunculidae). Galathea 11:119-124.

Dunk, K. v. d., and P. Lauterer. 1998. More details on the rare Eudorylas furvulus Collin and Microcephalops opacus (Fallen) comb. n. (Diptera: Pipunculidae). Acta Musei Moraviae - Scientiae Biologicae 82:163-171.

Földvári, M. 2003a. Contributions to the taxonomy of Afrotropical Eudorylini (Diptera, Pipunculidae). Folia Entomologica Hungarica. Rovartani Közlemények 64:267-276.

Földvári, M. 2003b. New Afrotropical species from the tribe Eudorylini (Diptera, Pipunculidae). Annales Historico-Naturales Musei Nationalis Hungarici 95:161-171.

Hardy, D. E. 1943. A revision of Nearctic Dorilaidae (Pipunculidae). University of Kansas Science Bulletin 29:1-231.

Huq, S. B. 1985. Über Eudorylas subterminalis Coll. (Dipt., Pipunculidae) als parasit bei Cicadellidae (Homoptera) in Berlin (West) [Studies on Eudorylas subterminalis (Diptera, Pipunculidae) parasitizing some Cicadellidae (Homoptera) in Berlin (West)]. Anzeiger fur Schädlingskunde, Pflanzenschutz, Umweltschutz 58:147-149.

Huq, S. B. 1986a. Description of final instar of Eudorylas subterminalis Collin (Diptera: Pipunculidae), an endoparasite of some leafhoppers (Homoptera : Cicadellidae). Bangladesh Journal of Zoology 14:19-23.

Huq, S. B. 1986b. Studies on the mating behaviour including some biological aspects of Eudorylas subterminalis Collin (Diptera: Pipunculidae), endoparasite of cicadellid leafhoppers (Homoptera: Cicadellidae). Zeitschrift für Angewandte Zoologie 73:281-292.

International Commision on Zoological Nomenclature. 2002. Eudorylas Aczél, 1940 and Microcephalops De Meyer, 1989 (Insecta, Diptera): conserved by the designation of Pipunculus fuscipes Zetterstedt, 1844 as the type species of Eudorylas. Bulletin of Zoological Nomenclature 59:143-144.

International Commission on Zoological Nomenclature. 2002. Opinion 2000 (Case 3132). Eudorylas Aczél, 1940 and Microcephalops De Meyer, 1989 (Insecta, Diptera): conserved by the designation of Pipunculus fuscipes Zetterstedt, 1844 as the type species of Eudorylas. Bulletin of Zoological Nomenclature 59:143-144.

Irwin, A. G. 1983. Eudorylas terminalis (Thom.) (Dipt., Pipunculidae) in Norfolk and Argyll. The Entomologist's monthly Magazine 119:80.

Kapoor, V. C., J. S. Grewal, and S. K. Sharma. 1987. Indian Pipunculids (Diptera: Pipunculidae). A Comprehensive Monograph. Atlantic Publishers & Distributors, New Delhi, India.

Kehlmaier, C. 2001. Records of Stratiomyidae, Pipunculidae and Conopidae (Diptera) from northern Spain. Munibe (Ciencias Naturales - Natur Zientziak) 51:79-84.

Kehlmaier, C. 2005. Taxonomic revision of European Eudorylini (Insecta, Diptera, Pipunculidae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 41:45-353.

Kehlmaier, C. (2005): Taxonomic studies on Palaearctic and Oriental Eudorylini (Diptera: Pipunculidae), with the description of three new species.- Zootaxa 1030: 1-48; Brisbane.

Kim, J. P., and C. H. Kim. 1986. Parasitism of green rice leafhopper, Nephotettix cincticeps Uhler by Alloneura oryzaetora, Eudorylas cruciator and Eudorylas tsuboii (Pipunculidae: Diptera) in southern rice cultural area. Research Reports of the Rural Development Administration, Plant, Environment, Mycology and Farm Products Utilization, Korea Republic 28:82-85.

Kozánek, M. 1988. Description of two new species of the genus Eudorylas (Diptera, Pipunculidae) from Soviet Middle Asia. Biológia (Bratislava) 43:945-948.

Kozánek, M. 1993. Descriptions of five new Eudorylas Aczél species from Central Europe (Diptera: Pipunculidae). Entomological Problems 24:39-50.

Kozánek, M., and C. Kehlmaier. 2004. Pipunculidae of Slovakia: additions and corrections to faunal list, with a description of a new Eudorylas (Diptera). Entomological Problems 34:21-36.

Kozánek, M., and Y. J. Kwon. 1991. Classification of the Family Pipunculidae from Korea (Diptera) Part II. On the genus Eudorylas Aczel from North Korea. Korean Journal of Entomology 21:157-165.

Kozánek, M., S. J. Suh, and Y. J. Kwon. 2004. Taxonomic review of the genus Eudorylas Aczél (Diptera: Pipunculidae) in Korea. Korean Journal of Entomology 34:105-111.

Kuznetzov, S. Y. 1993a. A new Australian species of Eudorylas Aczél (Diptera, Pipunculidae). Dipterological Research 4:145-150.

Kuznetzov, S. Y. 1993b. A new species of Eudorylas Aczél (Diptera, Pipunculidae) from Turkmenistan [sic]. Dipterological Research 4:201-207.

Kuznetzov, S. Y. 1994a. A new species of Eudorylas Aczél (Diptera, Pipunculidae) from the Middle Asia. Dipterological Research 5:33-37.

Kuznetzov, S. Y. 1994b. Short notes on synonymy and nomenclature of Pipunculidae (Diptera). Dipterological Research 5:105.

Kuznetzov, S. Y. 1995a. Eudorylas electus (Hardy): New record and redescription (Diptera, Pipunculidae). Acta Zoologica Fennica 199:33-35.

Kuznetzov, S. Y. 1995b. Neodorylas gen.n., with a key to the world genera, new synonymy and notes on the higher classification of the Pipunculidae (Diptera). International Journal of Dipterological Research 6:321-333.

McLean, I. F. G. 1981. A third British record of Eudorylas terminalis (Thom.) (Dipt., Pipunculidae). The Entomologist's monthly Magazine 117:29.

Morakote, R., Y. Hirashima, and K. Yano. 1990. A systematic study of the Japanese Pipunculidae (Diptera) Part V. The genus Eudorylas Aczél. Journal of the Faculty of Agriculture, Kyushu University 34:281-355.

Rafael, J. A. 1985. Pipunculidae (Diptera) da regiăo Neotropical. II. Redescription and correction of the generic position of Pipunculus flavicornis Williston and P. xanthopodus Williston, formerly in the genus Allomethus Hardy. Revista Brasileira de Entomologia 29:549-553.

Rafael, J. A. 1989. Descriçăo de duas espécies novas de Eudorylas Aczél (Diptera: Pipunculidae) de floresta inundável e de terra firme nas proximidades de Manaus, Amazonas, Brasil. Amazoniana 10:401-405.

Rafael, J. A. 1991. Espécies de Eudorylas Aczél do Peru (Diptera: Pipunculidae). Boletim do Museu Paraense Emilio Goeldi, Séries Zoologia 7:151-161.

Rafael, J. A. 1993. Espécies de Eudorylas Aczél (Diptera, Pipunculidae) do México e América Central. Revista Brasileira de Entomologia 37:751-762.

Rafael, J. A. 1995. Espécies de Eudorylas Aczél (Diptera, Pipunculidae) da América do Sul. Revista Brasileira de Entomologia 39:793-838.

Rafael, J. A. 1996. Pipunculidae (Insecta, Diptera) of the Dominican Republic: new records and description of new species. Annals of the Carnegie Museum 65:363-381.

Rafael, J. A., and R. Ale-Rocha. 1997. Chilean Pipunculidae (Diptera). Key to genera, new records, description of a new species of Eudorylas Aczél and a catalog of species. Revista Chilena de Entomologia 24:31-36.

Rafael, J. A., and M. D. d. S. Menezes. 1999. Taxonomic review of Costa Rican Pipunculidae (Insecta: Diptera). Revista de biologia tropical 47:513-534.

Rafael, J. A., and M. d. S. d. S. Rosa. 1989. Novas espécies de Eudorylas Aczél da Estaçăo Ecológica de Maracá, Roraima, Brasil (Diptera: Pipunculidae). Memorias do Instituto Oswaldo Cruz Rio De Janeiro 84:449-454.

Rafael, J. A., and M. S. S. Rosa. 1991. Pipunculidae (Diptera) da Estaçăo Ecológica de Maracá e da localidade de Pacaraima, Roraima, Brasil. Acta Amazonica 21:337-350.

Skevington, J. H. 2003. Revision of Australian Eudorylas Aczél (Diptera, Pipunculidae). Studia dipterologica 9:621-672.

Skevington, J., and S. A. Marshall. 1997. First record of a big-headed fly, Eudorylas alternatus (Cresson) (Diptera: Pipunculidae), reared from the subfamily Cicadellinae (Homoptera: Cicadellidae), with an overview of pipunculid-host associations in the Nearctic Region. The Canadian Entomologist 129:387-398.

Skevington, J. H., and D. K. Yeates. 2001. Phylogenetic classification of Eudorylini (Diptera: Pipunculidae). Systematic Entomology 26:421-452.

Vega, F. E., P. Barbosa, and A. P. Panduro. 1991. Eudorylas (Metadorylas) sp. (Diptera: Pipunculidae): a previously unreported parasitoid of Dalbulus maidis (Delong and Wolcott) and Dalbulus elimatus (Ball) (Homoptera: Cicadellidae). The Canadian Entomologist 123:241-242.

Title Illustrations
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Scientific Name Eudorylas
Specimen Condition Live Specimen
Sex Male
Copyright © Steve Marshall
Scientific Name Eudorylas mutillatus Loew
Identified By Jeff Skevington
Sex Female
Life Cycle Stage adult
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About This Page

Agriculture and Agri-Food Canada, Ottawa, Ontario, Canada

Correspondence regarding this page should be directed to Jeff Skevington at

Page: Tree of Life Eudorylas. Authored by Jeff Skevington. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Skevington, Jeff. 2005. Eudorylas. Version 02 November 2005 (under construction). http://tolweb.org/Eudorylas/54653/2005.11.02 in The Tree of Life Web Project, http://tolweb.org/

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